Adaptive advantages of dietary mixing different-aged foliage within conifers for a generalist defoliator. 2015. Johns, R.C.; Tobita, H.; Hara, H.; Ozaki, K. Ecological Research 30:783-802.
Dietary mixing within the crown of a decidous conifer enhances the fitness of a specialist sawfly. 2012. R. Johns; K. Ozaki; H. Tobita. Animal Behaviour 84: 1393-1400.